Observation 140595: Amanita homolae Tulloss nom. prov.

Solitary, found in mixed forest of birch spp, red oak, spruce-pine-hemlock. Cap with striate margin; entirely white, no gray on disc when fresh but dried is a bit gray. Saccate volva white and pretty large, not constricted or flimsy or crumbling like Amanita borealisorora; exannulate.

Proposed Names

31% (2)
Used references: Rod Tulloss’s Amanitaceae website
27% (1)
Used references: Amanitaceae website of RET

Please login to propose your own names and vote on existing names.

Eye3 = Observer’s choice
Eyes3 = Current consensus


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Thanks Britt,
By: groundhog
2013-10-29 12:19:14 PDT (-0700)

This material has been recieved and accessioned into Rod’s herbarium. We have scheduled it for DNA sequencing.
-Naomi (working with RET)

Thank you, Britt. Also, with a long “aside” regarding citrinoid taxa.
By: R. E. Tulloss (ret)
2013-09-23 15:02:24 PDT (-0700)

A couple of these have appeared on MO this years. There may be some projects coming up through which I may have access to sequencing in section Vaginatae. It will be valuable to have recent collections if these can be linked to the species concepts that I’ve proposed over the years on the Amanitaceae Studies website.


By the way, are you seeing citrina-like material now In Wisconsin? You may have heard of the citrinoid amanitas project that is based in Dr. Karen Hughes’ work at the University of Tennessee. I’ve been involved in that for a little while now, and three citrinoid taxa have been segregated in eastern North America. One of these appears to be A. brunnescens var. straminea, which I’ve thought for some time was not brunnescens. At any rate, we don’t have much material from the midwestern states of the U.S….even though the type of brunnescens var. straminea comes from Michigan, it dates back to the 1930s and is probably a forensic project if DNA can be extracted from it at all. We can separate the three taxa by spore size and shape as well as genetically (at least, it looks to us as though we can do that). At any rate, a set of samples of citrinoids from Wisconsin would be very helpful. Anybody reading this from Michigan? If so, PLEASE, help us out with samples from the home state of brunnescens var. straminea. In New Jersey, all three of the citrinoids are present in oak-conifer forest. More than once, all three have come in from a single foray carried out in a relatively small area such as the periphery of a smallish pond.

Here’s a suggestion for taking multiple collections from a single site:
1. Separate collections by the various shades of yellow in the cap. It’s OK to make subtle distinctions
2. Within the above separated groups, separate material by the size of the bulb.
3. Don’t worry about whether the material has purple, lavender or amethyst stains (all three taxa produce this reaction!).

Submit one collection each from the collections separated by the two stage method.

If you wish to try separation by spore shape, a comparison of the sporographs (graphs plotting size and shape simultaneously) for the three genetically distinct taxa is presented near the bottom of the technical tab on this page:


Here’s what we think we know about the three genetically distinguishable taxa:

1. Amanita sp-lavendula01 is an un-named infraspecific hybrid with multiple, different nrITS (barcode) gene copies in a single fruiting body. [The second article about different sorts of hybrids in agarics by Dr. Hughes is available as a prepublication format PDF on the Mycologia website. It is scheduled for publication…probably in the next issue. The citrinoid hybrid is mistakenly called Amanita citrina f. lavendula in the article. We’ll fix that over the next two years. The point is that this hybrid is a wild and crazy gal.]

2. Amanita sp-lavendula02 is Amanita citrina f. lavendula by comparison with possible lectotypes from Coker’s herbarium at NCU (University of North Carolina, Chapel Hill).

3. Amanita sp-lavendul03 is mentioned above and is A. brunnescens var. straminea by comparison with the remaining isotype of that species in MICH. The holotype was lost with E.-J. Gilbert’s herbarium, which was never deposited in P (Paris) as planned.

So there’s the story. Except for Missouri, we have very little coverage from west of the Mississippi. So citrinoids from the Great Plains are also solicited.

A long side trip from homolae, but important because 2013 is our main year for accumulation of collections for the citrinoid project.

Very best,


I’ll send it
By: Britt Bunyard (Fungi magazine) (bbunyard)
2013-09-23 13:46:13 PDT (-0700)

right out to you.

Do you have dried material from this observation?
By: R. E. Tulloss (ret)
2013-09-23 13:16:04 PDT (-0700)

If so, Britt, I’d be interested in seeing it.

Very best,


Hello, Britt.
By: R. E. Tulloss (ret)
2013-07-23 07:56:41 PDT (-0700)

Amanita rasitabula is very pure white and the volva is not like what you’ve described here in terms of robustness. The rasitabula volva is rather flimsy and is often partially fragmented and barely holding together. Your material somewhat suggests A. homolae to me. This is a mushroom I know from Maine and might occur in the north central states as well. ??

Very best,


Hi Rocky…
By: Britt Bunyard (Fungi magazine) (bbunyard)
2013-07-23 03:54:07 PDT (-0700)

My specimen had no gray on it whatsoever, but was only a single specimen.

By: Rocky Houghtby
2013-07-22 20:01:40 PDT (-0700)

I think this is the same species:


I collected it for two weeks exclusively beneath Red Oak.

It’s quite probable that people would have misapplied…
By: R. E. Tulloss (ret)
2013-07-22 09:41:13 PDT (-0700)

vaginata var. alba to this mushroom; however, I think this is different from the species to which the European name has been applied along the east coast of the U.S. (A. rasitabula).




Very best,


Created: 2013-07-22 06:53:04 PDT (-0700)
Last modified: 2018-01-04 06:20:34 PST (-0800)
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