Observation 54305: Umbilicaria vellea (L.) Hoffm.
When: 2010-09-15
Herbarium specimen reported

Notes: On outcrop on narrow ridge just above pass (to north) with several other species of Umbilicaria, Pseudephebe, etc.

20100915.106

Images

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close-up showing two kinds of rhizines clearly
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ball-tipped or knobbed black rhizines
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slender, pale, branched rhizines

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Bruce Ryan’s glossary
By: Chris Parrish (kitparrish)
2010-10-01 19:13:28 PDT (-0700)

Two entries in Bruce Ryan’s glossary:

THALLOSPORE a vegetative (asexual) propagule, granular in appearance, composed of one to several pigmented, thick walled fungal cells, borne on the underside or edges of lobate to umbilicate thalli; the term is also applied in other ways (at least in nonlichenized fungi).

RHIZINOMORPH rhizinelike organ not attaching thallus to its substrate, in peltate lichens (e.g., some species of Umbilicaria and Dermatocarpon)

fascinating structures
By: Chris Parrish (kitparrish)
2010-10-01 19:10:12 PDT (-0700)

Bruce Ryan’s notes on Umbilicaria are excellent … much more detailed than the other references I have checked. In particular, read through his key to the “U. vellea group sensu lato,” analizing seven different species. He also gives strong hints as to the biological functions of these strange structures on the undersides of non-fertile, non-sorediate, non-isidiate Umbilicaria (involving minute black granules called “thallospores”), and that may indicate why he is using the term “rhizinomorphs” (because their function has to do with these thallospores, rather than connecting to the substrate as for conventional rhizines).

Created: 2010-10-01 17:50:00 PDT (-0700)
Last modified: 2010-10-01 22:28:49 PDT (-0700)
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