Pileus 4.3-1 1 × 1.5-3 cm; pale charcoal, “li- noleum brown,” “camel,” “sunburn,” “choco-late,” “teak,” “brownish grey” (5E7; 6D-F4,5: 8E,F2-4), convex, non-staining or darker upon injury, non-staining or darker in ammonia, even, velvety, dry, occasionally convoluted, infre-quently cracked or areolate. Pileus context 0.5- 2.5 cm thick, gray to pallid gray, staining “peach,” “reddish orange” (7A4,5; 7B6) and finally fus- cous upon injury. Tubes 0.3-1.1 cm long, gray, staining orange, “greyish red” (7,8B6) and finally dark fuscous when injured, depressed about the stipe; tube mouths small (I-3/mm), black when young, gray in age, staining grayish orange and finally “brownish grey” (8C,D2-3). Stipe 4.5-8.5 × 1-3.5 cm, obclavate, subobclavate with a basal taper, blackish brown, charcoal, “brownish grey” (8E,F2,3; 6F2,3), lighter above, darker, exan-nulate; pruinose overall and coarsely reticulate over the upper third to halc context solid, pallid gray to dark gray, staining orange to red-orange, “greyish red” (7B6) and finally dark fuscous. Ba- sidiospores “brownish orange,” “Titian red,” “terra cotta,” “greyish red,” "brownish red (7C4; 7D6,7; 8C,D4-6) in deposit. Pileus cuticle a palisade trichoderm; terminal cells (FIG. 1) 19.5-65 × 5.2-1 1.7 ym (a = 39.8 × 8.3 ym) cylindric, opaque, cream, cinnamon- gold, smoky tan in KOH, light gold to dark gold- brown in Melzer’s reagent, blue to blue-green in cotton blue. Tube tramal hyphae 3.9-12.3 ym diam (ii= 7.4 wm), bilateral-divergent (boletoid), hyaline in KOH, Melzer’s reagent, and cotton blue; walls occasionally with circumferential thickenings. Oleiferous hyphae of the tube trama 2.6-13 pm diam (X = 7.2 pm), gold to dark cin- namon-tan in KOH, dark rust-brown to dark brown in Melzer’s reagent, blue to blue-green in cotton blue; occasionally with circumferential thickenings. Clamp connections absent. Basidia (FIG. 2) 18.2-39 × 6.5-1 1.7 pm (f = 27.6 × 8.6 pm) clavate to narrowly clavate, opaque cream in KOH, hyaline to opaque yellow in Melzer’s reagent, blue in cotton blue. Pleurocystidia (FIG. 3) 27.3-85.8 × 7.8-14.3 Mm (X = 52.9 × 10.5 . . pm), fusoid-ventricose, ventricose-rostrate, broadly lanceolate, hyaline in KOH, Melzer’s re- agent, and cotton blue; proximal walls occasion- ally thickened. Pseudocystidia (FIG. 4) 26-80.6 × 6.5-15.6 pm (R = 43.6 × 9.4 pm), most fre- quent near tube mouths, fusoid-ventricose, nar- rowly fusoid-ventricose, rarely cylindric near tube mouths, cinnamon, cinnamon-gold, or brown (near tube mouths) in KOH, dark cinnamon, sor- did brown, red-brown, or olive-gold in Melzer’s reagent, blue to blue-green in cotton blue; con- tents granular to coagulate. Cheilocystidia (FIG. 5) 39-1 14.1 × 5.2-1 1.7 pm (f = 72.3 × 7.8 pm), infrequent to abundant, narrowly fusoid-ventri- cose, broadly lanceolate, lanceolate, occasionally with secondary basal septum; and attenuate dis- tal end, pale cream to pale cinnamon in KOH, pale cinnamon to dark cinnamon-gold in Mel- zer’s reagent; contents pale blue to blue-green in cotton blue, distally coagulate. Stipe dermato- cystidia (FIG. 6) 18.2-65 × 6.5-15.6 pm (f = 39.1 × 1 1.2 pm), fusoid-ventricose, clavate, cy- lindric, cinnamon-brown (walls darker) to pale tan in KOH, dark brown to gold-brown in Mel- zer’s reagent, blue to blue-green in cotton blue; proximal wall infrequently thickened, rarely with secondary septa. Basidiospores (FIG. 7) 7.8-14.3 × 3-5.2 pm (X = 11.5 × 4 pm), Q(E) (length/ width range) = 2.3-3.7 (f = 2.9), inequilateral in profile view, fusiform-elliptic to elliptic in face view, yellow-cream to opaque yellow-cream in KOH, opaque yellow to dark yellow-tan in Mel- zer’s reagent, hyaline in cotton blue; surface smooth, distal end acute; walls continuous. Macrochemistry. Pileus surface vinaceous lack in KOH; pileus context vinaceous pink in KOH; stipe context red-orange in KOH.
Habitat and distribution. Tylopilus griseocarneus occurs in sandy soils under Quercus and Pinus species in the Atlantic and Gulf Coastal Plain forests of southern New Jersey and south- ern Louisiana. We expect its distribution to be continuous throughout the Coastal Plain be-tween Louisiana and New Jersey.