When: 2008-04-12

Collection location: Sebastopol, Sonoma Co., California, USA [Click for map]

Who: Darvin DeShazer (darv)

Specimen available

Notes:
Collected and dried 28 fruiting bodies. They were growing in damp, shady soil under Rubus and Corylus cornuta with moss and low growing plants. Spore print was white. Spores 9-10 µm
Collection # DD636

There might be two names for this fungus. It seems to be a perfect fit for the keys and the descriptions by both Bigelow and Gregory. Redhead dismisses all species in Bigelow, but fails to state why this one does not match. When I asked Denise Gregory about this species she wrote “Hi Darvin, I don’t believe that Clitocybe scabriuscula is truly a Clitocybe, even though I have it listed in my monograph.” After doing a literature search it seems that Clitocybe scabriuscula has NOT been moved to Omphalina, Contumyces or anything else. It currently remains Clitocybe scabriuscula on both Zip Code Zoo and Index Fungorum and does NOT appear in any Omphalina key, including the lengthy keys in Moser (33 species) or Hanson (27 species). Bigelow keys out at least 45 species of omphalinoid Clitocybes in his book but does NOT include Omphalina rosella.
On the other hand, the two color photos in Antonin and in Dahncke are an excellent fit. The description in Redhead, supported by the opinion of Dr. Desjardin, strongly supports Contumyces rosellus var. vinacea.
Following each reference is a germane comment for this collection.

Species Lists

Images

Young, damp cap unexpanded.
Cap starting to dry out.
Young cap drying out.
Mature cap with granules of pink pigments.
Mature cap drying out and the margin turning white.
Six mature caps with the margin drying out to white.
Gills white.
Gills subdistant and thick, margin white when young and damp.
Gills with insect larvae.
Gills starting to turn pinkish.
Gills turning pinkish and a pruinose apex on the stipe.
Marginate gills with pinkish color.
Gill attachment to the stipe.
On the stipe are clusters of cystidia and granules of reddish pigments.
Best seen when viewed on FULL SIZE.
Lateral and terminal pileocystidia projecting from the pileipellis.
Two photos in a row taken with light and dark field to show the same clamp connections on the hyphae of the pileipellis.
Two photos in a row taken with light and dark field to show the same clamp connections on the hyphae of the pileipellis.
Fascicles of caulocystidia on the stipe. Image taken with phase objective at 200X to show the 3-D relief of the hyphae.

Proposed Names

92% (2)
Recognized by sight: Color photo online
Used references:
Antonín, Vladimír and Machiel E. Noordeloos. 2004. A Monograph Of The Genera Hemimycena, Delicatula, Fayodia, Gamundia, Myxomphalia, Resinomycena, Rickenella, and Xeromphalina (Tribus Mycenae sensu Singer, Mycena excluded) In Europe. IHW Verlag, Eching. 279p. (Color photo of Contumyces rosella and full description with micodrawings; only three species listed)
Based on microscopic features: Spores 9-10 µm
Clamps found almost everywhere.

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= Observer’s choice
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